Wrilii was originally a Zo-Matoran of the Tren Krom Peninsula. He lived in Zo-Koro near the Ojrax River. He worked there as a navigator and made very close friends with a Ga-Matoran named Coryna, a sailor from a village nearby. He eventually made rivals with a criminal Ta-Matoran named Juzto. After the latter made a successful raid on Wrilii's village with several others, he got into a battle with the Zo-Matoran. Wrilii managed to defeat and capture all of the Matoran outlaws save Juzto.
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