Mogilevsky was born to Jewish parents in Pavlograd. In 1903 he joined the RSDRP, and in 1904 Mogilevsky was arrested. In 1906 he left Russia for Switzerland where Mogilevsky met Vladimir Lenin who recommended him to be admitted into Bolshevik section of the party. At the outbreak of the World War I he returned to Russia and was a soldier on the front. Mogilevsky participated in the 1917 October Revolution. During the Russian Civil War he was appointed to various positions in the GPU. Since May 1922 Mogilevsky headed the Caucausian GPU and was responsible for intelligence in Iran and Turkey. He participated in stifling the August Uprising in Georgian SSR. Mogilevsky was decorated with the Order of the Red Banner in 1924 for his outstanding activities in the suppression of the uprising.
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