12,000 years after their last clash with mankind, the Shadow Angels return to attack the human cities and kidnap its inhabitants. Looking for the reincarnation of Apollonius, a Shadow Angel who turned against his own kind to protect the humans, Silvia de Alisia has an encounter with Apollo, a reckless and animalistic orphan from Arc City who somehow manages to unlock the true potential of mankind's greatest weapon, the Mechanical Angel Aquarion.
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