If the introduction is anything more than Gwyn's 'Official History' (the existence of the Everlasting whelp and the Archtrees is evidence that it is), the world originally belonged not to the Darkness but to the Stone Dragons and 'humans' were scraggly... things until four of them managed to grab the Souls of Lords (from where is not explained either). The condition of humans at the beginning of the game proper is partly a product of the Age of Fire and any magical effects thereof, so once the Souls of Lords and the First Flame are extinguished the result should be an Age of Undead rather than an age of Humans -- unless there's some unrevealed original source of Humanity besides other humans and monsters (which as one WMG suggest may have gotten it from human victims).
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