abstract
| - The genus Gansus contains a single species, G. yumenensis,[2] which was about the size of a pigeon and similar in appearance to loons and diving ducks.[5] It had many features common among modern birds, and also retained some primitive traits such as its clawed wings.[5] Gansus was discovered in the form of a single fossil foot in 1981. Five more well-preserved fossils were found in 2003–2004 in mudstone at the site of an ancient lake at Changma, Gansu; the geological stratum in which the fossils were found is the Xiagou Formation. Their bodies had settled in anoxic mud and were soon covered with further extremely fine silty sediments. Without oxygen, their remains resisted decay: these specimens preserved remains of flight feathers and traces of the webbing between their toes.[5] In 2011, there were described nine additional specimens, that supported on the basis of a statistic analysis of the sternum and elements of the legs, the hypothesis that Gansus was a volant bird.[6] You et al. (2006) concluded that the anatomical characteristics of Gansus were similar to foot-propelled diving birds, such as Hesperornis (from the Cretaceous) and the loons (Gaviidae) and grebes (Podicipedidae).[5] On the other hand, Li et al. (2011) concluded that Gansus showed a more similar morphology to ducks.[6] Two years later, Nudds et al. (2013) showed that the pectoral limb length proportions of Gansus were most similar to swifts and hummingbirds (Apodiformes), while the pelvic limb length proportions fell within the modern birds (Neornithes), showing similarities with grebes (Podicipedidae), albatross (Diomedeidae) and cormorants (Phalacrocoracidae), suggesting that Gansus was both volant and capable of diving to some degree using either foot-propelled or, perhaps, both its wings and its feet for underwater locomotion.
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