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Hepburn, H.R. 1978a. The bee louse. South African Bee Journal 50(6): 11-12. Hepburn, H. R. 1978b. Some honeybee research activities at the University of the Witwatersrand. South African Bee Journal 50(5): 11-14. Hepurn, H. R. 1979a. Braula coeca. South African Bee Journal 50(6): 11-12. Hepurn, H. R. 1979b. Pollination versus honey: the beekeeper’s seesaw. South African Bee Journal 51(2): 3-7. Hepburn, H. R. 1980. How honeybees measure their pollen loads. South African Bee Journal 52(3): 11-13. Hepurn, H. R. 1981a. Efficacy of pollen traps. South African Bee Journal 53(5): 8-9. Hepurn, H. R. 1981b. Fruit tree pollination. South African Bee Journal 53(5): 14-15. Hepurn, H. R. 1981c. Is the Langstroth brood chamber deep enough for African bees? South African Bee Journal 53(4): 9-11. Hepurn, H. R. 1982a. Comb production and the new foundation. South African Bee Journal 54: 56-62. Hepurn, H. R. 1982b. The bee scorpion. South African Bee Journal 54(2): 39-41. Hepburn, H. R. 1983a. 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Journal of the Entomological Society of Southern Africa 54(1): 84-85. Hepurn, H. R. 1991b. Some apicultural puzzles. South African Bee Journal 63: 13-15. Hepurn, H. R. 1991c. The unique Cape bee and some apicultural puzzles. Beeline 28: 12-15. Hepurn, H. R. 1992a. “Follow-flow” brood rearing in capensis and scutellata. South African Bee Journal 64: 104-108. Hepurn, H. R. 1992b. Drongos: honey guides of the eastern Cape. South African Bee Journal 64: 40-41. Hepurn, H. R. 1992c. Pheromonal and ovarial development covary in Cape worker honeybees, Apis mellifera capensis. Naturwissenschaften 79: 523-524. Hepurn, H. R. 1992d. The biology of queen production in the Cape honeybee. South African Bee Journal 64: 132-138. Hepurn, H. R. 1993a. Heuning tydens die Tweede Vryheidsoorlog. South African Bee Journal 65: 10-12. Hepurn, H. R. 1993b. Swarming, absconding and migration in southern African bees. South African Bee Journal 65: 61-66. Hepburn, H. R. 1994. Reproductive cycling and hierarchical competition in Cape honeybees, Apis mellifera capensis Esch. Apidologie 25: 38-48. Hepurn, H. R. 1995. Flight and foraging in capensis and scutellata. South African Bee Journal 67: 38-42. Hepurn, H. R. 1998. Reciprocal interactions between honeybees and combs in the integration of some colony functions in Apis mellifera. Apidologie 29: 47-66. Hepurn, H. R. 2000. Honey bee (Apis mellifera) classification and the confounding effects of trinomial nomenclature. Anais do Encontro sobre Abelhas 4: 188-196. Hepburn, H. R. 2001. The enigmatic Cape honey bee, Apis mellifera capensis. Bee World 82(4): 181-191. also in Honeybee Science 22: 49-56. [In Japanese]. Hepurn, H. R. 2002. Apis mellifera capensis: concept and reality. South African Bee Journal 74(2): 37-39, 53-62. Hepurn, H. R. & M. H. Allsopp, 1994. Reproductive conflict between honeybees: usurpation of Apis mellifera scutellata colonies by Apis mellifera capensis. South African Journal of Science 90: 247-249. Hepurn, H. R., E. Armstrong & S. P. Kurstjens, 1983. The ductility of native beeswax is optimally related to honeybee colony temperature. South African Journal of Science 79: 416-417. Hepurn, H. R., R. T. F. Bernard, B. C. Davidson, W. J. Muller, P. Lloyd, S. P. Kurstjens & S. L. Vincent, 1991. Synthesis and secretion of beeswax in honeybees. Apidologie 22: 21-36. Hepburn, H. R., R. C. Cantrill, P. R. Thompson & E. Kennedi, 1979. Metabolism of carbohydrate, lipid and protein during development of sealed worker brood of the African honeybee Apis mellifera adansonii. Journal of Apicultural Research 18(1): 30-35. Hepburn, H. R. & H. D. Chandler, 1975. A new classification for arthropod cuticles. South African Journal of Science 71(10): 311-312. Hepburn, H. R. & H. D. Chandler, 1976a. Material properties of arthropod cuticles: the arthrodial membranes. Journal of Comparative Physiology 109(2): 177-198. Hepburn, H. R. & H. D. Chandler, 1976b. Mechanical hysteresis of insect cuticles. Journal of Insect Physiology 22(2): 221-226. Hepburn, H. R., H. D. Chandler & M. R. Davidoff, 1979. Extensometric properties of insect fibroins: the green lacewing cross-beta, honeybee alpha-helical and greater waxmoth parallel-beta conformations [Chrysopa carnea, Apis mellifera adansonii, Galleria mellonella]. Insect Biochemistry 9(1): 69-77. Hepburn, H. R. & R. M. Crewe, 1990. Defining the Cape South Africa honey bee reproductive traits of queenless workers. South African Journal of Science 86: 524-527. Hepburn, H. R. & R. M. Crewe, 1991a. Portrait of the Cape honeybee, Apis mellifera capensis. Apidologie 22(6): 567-580. Hepurn, H. R. & R. M. Crewe, 1991b. Geography of the Cape honeybee based on laying worker performance. South African Bee Journal 63: 51-59. Hepburn, H. R. & S. G. Guye, 1993. An annotated bibliography of the Cape honeybee, Apis mellifera capensis Eschscholtz (Hymenoptera: Apidae). African Entomology 1(2): 235-252. Hepburn, H. R. & C. Hepburn, 2005. Bibliography of Apis florae. Apidologie 36(3): 377-378. Hepburn, H. R., J. J. Hugo, D. Mitchell, M. J. M. Nijland & A. G. Scrimgeour, 1984. On the energetic costs of wax production by the African honeybee, Apis mellifera adansonii. South African Journal of Science 80(8): 363-368. Hepurn, H. R. & Guillarmod A. Jacot, 1991. The Cape honeybee and the fynbos biome. South African Journal of Science 87: 70-73. Hepurn, H. R., G. E. Jones & R. Kirby, 1994. Introgression between Apis mellifera capensis Escholtz and Apis mellifera scutellata Lepeletier: the sting pheromones. Apidologie 25: 557-565. Hepburn, H. R. & S. P. Kurstjens, 1984. On the strength of propolis (bee glue). Naturwissenschaften 71(11): 591-592. Hepburn, H. R. & S. P. Kurstjens, 1988. The combs of the honeybee as composite materials. Apidologie 19: 25-36. Hepburn, H. R. & P. C. Magnuson, 1988. Nectar storage in relation to wax secretion by honeybees. Journal of Apicultural Research 27(2): 90-92. Hepburn, H. R., P. Magnuson, L. Herbert & L. A. Whiffler, 1991. The development of laying workers in field collected colonies of the Cape honeybee. J. Apic. Res. 30: 107-112. Hepurn, H. R. & G. Mitchell, 1981. Milk and Honey, Essays in Agriculture. Elsevier/North Holland, Amsterdam 108 pp. Hepurn, H. R. & W. J. Muller, 1988. Wax secretion in honeybees. Thermal integrity of the festoon. Naturwissenschaften 75: 628-629. Hepurn, H. R., R. J. C. Nefdt & L. A. Whiffler, 1988. Queen loss in the Cape honeybee: the interactions of brood, laying workers (false queens) and queen cells. South African Journal of Science 84: 778-780. Hepburn, H.R. & S. E. Radloff, 1995. First approximation to a phenology of the honeybees (Apis mellifera) and flora of Africa. Oecologia 101(3): 265-273. Hepurn, H. R. & S. E. Radloff, 1996a. Beeswax exports and rainfall in the savanna woodlands of east central Africa. Apidologie 27: 473-478. Hepurn, H. R. & S. E. Radloff, 1996b. Population structure of the honeybees along a transect from the Sahara to the Pyrenees. Apidologie 27: 35-45. Hepurn, H. R. & S. E. Radloff, 1997. Biogeographical correlates of population variance in the honeybees (Apis mellifera) of Africa. Apidologie 28: 243-258. Hepurn, H. R. & S. E. Radloff, 1998. Honeybees of Africa. Springer-Verlag, Berlin 370 pp. Hepburn, H. R. & S. E. Radloff, 2002. Apis mellifera capensis: an essay on the subspecific classification of honeybees. Apidologie 33(2): 105-128. Hepburn, H. R., S. E. Radloff & S. Fuchs, 1998a. Population structure and the interface between Apis mellifera capensis and Apis mellifera scutellata. Apidologie 29: 333-346. Hepurn, H. R., S. E. Radloff & S. Fuchs, 1998b. Ecological and morphological differentiation of the honeybees, Apis mellifera Linnaeus (Hymenoptera: Apidae) of west Africa. African Entomology 6: 17-23. Hepburn, H. R., S. E. Radloff & S. Fuchs, 1999. Flight machinery dimensions of honeybees, Apis mellifera. Journal of comparative physiology. B. Biochemical, systemic, and environmental Physiology 169(2): 107-112. Hepurn, H. R., S. E. Radloff, S. Fuchs, G. W. Otis, S. Hadisoesilo & A. C. Tilde, 2003. Mutlivariate morphometric analysis of the Apis cerana populations of oceanic southern Asia. Apidologie [In Press]. Hepburn, H. R., S. E. Radloff & S. Oghiakhe, 2000. Mountain honeybees of Africa. Apidologie 31(2): 205-222. Hepburn, H. R., S. E. Radloff, G. W. Otis, S. Fuchs, L. R. Verma, T. Ken, T. Chaiyawong, G. Tahmasebi, R. Ebadi & S. Wongsiri, 2005. Apis florea: morphometrics, classification and biogeography. Apidologie 36(3): 359-376. Hepburn, H. R., S. E. Radloff, G. R. Steele & R. E. Brown, 1998. Dimensional aspects of flight in the honey bees of Africa. Journal of Apicultural Research 37(3): 147-154. Hepburn, H. R., S. E. Radloff, S. Verma & L. R. Verma, 2001. Morphometric analysis of Apis cerana populations in the southern Himalayan region. Apidologie 32: 1-13. Hepurn, H. R., S. L. Reece, P. Neumann, R F. A. Moritz & S. E. Radloff, 1999. Absconding in honeybees (Apis mellifera) in relation to queen status and mode of worker reproduction. Insectes Sociaux 46: 323-326. Hepburn, H. R. & J. R. Rigby, 1981a. Strength of beeswax foundation in relation to cell orientation. Journal of Apicultural Research 20(4): 234-238. Hepurn, H. R. & J. R. Rigby, 1981b. Beeswax candles from a silicone rubber mould. South African Bee Journal 53(4): 11-13. Hepburn, H. R. & D. C. Roberts, 1975. Stiffness and tanning of sclerites. J. Insect Physiol. 21(11): 1741-1746. Hepburn, H. R. & G. N. Sinclair, 1977. On the occurrence of a fungus disease in the compound eyes of the African drone. South African Bee Journal 49(6): 2-4. Hepburn, H. R., D. R. Smith, S. E. Radloff & G. W. Otis, 2001. Infraspecific categories of Apis cerana: morphometric, allozymal and mtDNA diversity. Apidologie 32(1): 3-23. Hepburn, H. R., M. H. Villet, G. E. Jones, A. R. Carter, V. I. Simon & W. Coetzer, 1993. Winter absconding as a dispersal mechanism of the Cape honeybee. South African Journal of Science 89(6): 294-297. Hepurn, H. R. & L. A. Whiffler, 1988. Dwarfs and queenballs in swarms of the Cape honeybee. South African Journal of Science 84: 851-852. Hepurn, H. R. & L. A. Whiffler, 1991. Construction defects define pattern and method in comb building by honeybees. Apidologie 22: 381-388. Hepburn, H. R., P. B. Wilson, H. D. Chandler & P. R. Thompson, 1978. Fractography of insect cuticle. Journal of Entomological Society of South Africa 41(1): 51-56. Hepurn, H. R., C. Youthed, P. Illgner, S. E. Radloff & R. E. Brown, 1998. Production of aerodynamic power in mountain honeybees (Apis mellifera). Naturwissenschaften 85: 389-390. Heppner, J. B. & D. H. Habeck, 1977. Insects associated with Polygonum (Polygonaceae) in north central Florida. II. Insects other than Lepidoptera. Fla. Entomol. 60: 167-170. Heras, C. & S. F. Gayubo, 1989. Bee fauna from the province of Zamora [Spain]: II. Megachilidae (Hymenoptera, Apoidea). Bol. Asoc. Esp. Entomol. 13: 237-250. Heraty, J. M., J. B. Woolley & D. C. Darling, 1994. Phylogenetic implications of the mesofurca and mesopostnotum in Hymenoptera. Journal of Hymenoptera Research 3: 241-277. Herbers, J. M. 1979. The evolution of sex-ratio strategies in hymenopteran societies. American Naturalist 114: 818-834. Herbert, E. W. Jr. 1979a. A new ash mixture for honeybees maintained on a synthetic diet. Journal of Apicultural Research 18(2): 144-147. Herbert, E. W. Jr. 1979b. Brood rearing by small caged honeybee colonies fed whey-yeast pollen substitutes. Journal of Apicultural Research 18(1): 43-46. Herbert, E. W., W. E. Bickley & H. Shimanuki, 1970. The brood-rearing capability of caged honey bees fed dandelion and mixed pollen diets. Journal of Economic Entomology 63(1): 215-218. Herbert, E. W. Jr., W. A. Bruce & H. Shimanuki, 1988. Control of Varroa jacobsoni on honey bees in packages using Apistan. American Bee Journal 128(9): 615-616.
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