The Tribe of Ancients were an advanced civilization appearing in Super Paper Mario that was led by four sages, including Merlimbis, a female sage that created the Pure Heart, Merloo, a male sage that wrote the Light Prognosticus to counteract the Dark Prognosticus, Merlight, another male sage that designed Flipside, and Merlumina, whose spirit guarded the Pure Heart in Yold Desert for 1,500 years. Their power and influence declined, however, after the Pixl Uprising.