Pterodactyloidea (derived from the Greek words πτερόν (pterón, for usual ptéryx) "wing", and δάκτυλος (dáctylos) "finger" meaning "winged finger", "wing-finger" or "finger-wing") forms one of the two suborders of pterosaurs ("wing lizards"), and contains the most advanced members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal rhamphorhynchoidea by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth. Many species had well developed crests on the skull, a form of display taken to extremes in giant-crested forms like Nyctosaurus and Tapejara. Pterodactyloids (specifically the family Azhdarchidae) were the last surviving pterosaurs when the order became extinct at the end of the Cretaceo