During the Great War, the majority of the Tonberries remained in their strongholds, the Temple of Uggalepih and Pso'Xja. However, a few elite members of the tribe were sent to the Shadow Lord to act as his secret agents. These Tonberries would often sneak into the castles of the allied forces and assassinate important generals or political figures. Feared around Vana'diel, these beastmen were often referred to as the "invisible killers". Nevertheless, according to Grav'iton, many Tonberries are actually warped Kuluu.