Spectre was a Guardian and a member of the Brotherhood of Guardians from Dark Mobius, a parallel future Zone. His history was presumably near identical to that of his counterpart from the Prime Zone, up until the point that Dark Enerjak took Spectre's descendant Knuckles as his newest host. Knuckles' aggressive efforts to bring his own version of order to Mobius led to conflict between him and the Brotherhood. In particular, Spectre's condemnation of Knuckles' association with the Dark Legion prompted the younger Echidna to send Spectre flying through a wall. (ASUB)