Dryopithecus was a genus of apes that is known from from Eastern Africa into Eurasia. It lived during the Upper Miocene period, from 12 to 9 million years ago, and probably includes the common ancestor of the lesser apes (gibbons and siamangs) and the great apes. The first species of Dryopithecus discovered was found in France in 1856. The five-cusp and fissure pattern of its molar teeth, known as the Y-5 arrangement, is typical of the dryopithecids and of hominoids in general. Other dryopithecids have been found in Hungary, Spain, and China.