Ankylosaur trackways were found in 1996 near Sucre, Bolivia, South America showing that these huge creatures could run at a "decent jog," according to Christian Meyer, a Swiss paleontologist working at the site. Speed estimates are made using the distance between the footprints, the animal's leg length, the pattern of the tracks, and other factors. Partial fossils have been found in England. Acanthopholis was named by the British biologist Thomas H. Huxley in 1865. Acanthopholis belonged to the: