The band's members are Twilight on (temporary) lead vocals, Rainbow Dash on electric guitar and (initially and later permanently) lead vocals, Applejack on bass, Rarity on keytar, Fluttershy on tambourine, Pinkie Pie on drums, and on one occasion as a guest, DJ Pon-3 on turntables. After Twilight leaves, Sunset Shimmer joins as rhythm guitarist and backup vocalist, and in Legend of Everfree, Twilight's human counterpart joins as a backup vocalist.