Jane-B337 was a recruit of the Special Proficiency, Assault, Reconnaissance and Tactical Augmented Nonconventional-force, Third Generation (SPARTAN-III) Program from Beta Company. Mid-training, she was one of several recruits taken away from the other trainees and trained into separate, secret two-man cells which, together, were known as The Headhunters. Upon graduation, Jane and her partner, Nessa-B256, went on missions behind enemy lines away from the rest of Beta Company, thus avoiding the catastrophic Battle at Pegasi Delta.