Milo is an evolved chimpanzee, he is the youngest son of the Ape King Caesar, and his Queen Cornelia, and the younger brother of Blue Eyes. Through his father, Milo is the youngest grandson of the late Bright Eyes, and the late Alpha, as well as the youngest adoptive grandson of the late Will Rodman, and the late Caroline Aranha, and the youngest adoptive great-grandson of the late Charles Rodman. He will be second-in-line to inherit the colony after his brother, Blue Eyes.