Titeuf was initially published in the fanzine Sauve qui peut ("Run for your lives") and noticed by Glénat executive Jean-Claude Camano. Zep joined Glénat in 1992 and Titeuf eventually became one of France's most popular comics. The first Titeuf book Dieu, le sexe et les bretelles (God, Sex And Suspenders) appeared in 1993 and sold only a few thousand copies, but the subsequent books gradually won over a colossal readership, and the series is now considered the greatest moneymaker in the French comics market. The series was adapted into an animated TV series in 2001, initially broadcast on Canal J. By 2008, Titeuf was the comic series with by far the largest publication in France, with over 1.8 million copies per year, three times the number of the second most popular series.