Of the four species, two (P. gunnari and P. klekowskii) are known from numerous remains found in Middle or Late Eocene strata (34 to 50 MYA) of the La Meseta Formation on Seymour Island, Antarctica. P. antarcticus, the first fossil penguin described, is only really known from a single incomplete tarsometatarsus found in the Late Oligocene Otekaike Limestone (23 to 28, possibly up to 34 MYA) at Kakanui, New Zealand, but numerous other bones have been tentatively assigned to the species. The other described New Zealand species, P. marplesi, is known from parts of a skeleton, mainly leg bones, from the Middle or Late Eocene Burnside Mudstone (34 to 40 MYA) at Burnside, Dunedin. To this species also a number of additional remains have been tentatively assigned. The problem with the indetermina