Zoro-Agruga occupied Heimdallr from S.371 to ca. S.470 and, with the dark breath it sprayed, turned the capital into the city of the dead. Zoro-Agruga's reign ended with the counter-attack of the Imperial family, led by Emperor Hector I, wielding the Vermillion Divine Knight Testa-Rossa. Upon its death, its blood claimed Hector's life and placed a curse on the Divine Knight. Its remains were locked away below the newly built Heimdallr.