The Late Devonian extinction was one of five major extinction events in the history of the Earth's biota. A major extinction occurred at the boundary that marks the beginning of the last phase of the Devonian period, the Famennian faunal stage, (the Frasnian-Famennian boundary), about 364 million years ago, when nearly all of the fossil agnathan fishes suddenly disappeared. A second strong pulse closed the Devonian period. Overall, 19% of all families and 50% of all genera went extinct.