Gran was an unjoined male Trill. The second child of Jayvin Vod and Audrid Dax, Gran was born in the 2250s, three years after his sister, Neema. Gran died in the year 2275, when he was in his mid-twenties. (DS9 - The Lives of Dax short story: "Sins of the Mother", TOS eBook: The More Things Change) Ezri Dax found that Brinner Finok reminded her of Gran, which made it difficult for her to pursue a romantic relationship with him. (DS9 episode: "Prodigal Daughter")