In oceanography and limnology, bioturbation is the displacement and mixing of sediment particles by benthic fauna (animals) or flora (plants). The mediators of bioturbation are typically annelid worms (e.g. polychaetes, oligochaetes), bivalves (e.g. mussels, clams), gastropods, holothurians, or any other infaunal or epifaunal organisms. Faunal activities, such as burrowing, ingestion and defecation of sediment grains, construction and maintenance of galleries, and infilling of abandoned dwellings, displace sediment grains and mix the sediment matrix. The sediment-water interface increases in area as a result of bioturbation, affecting chemical fluxes and thus exchange between the sediment and water column. Some organisms may further enhance chemical exchange by flushing their burrows with