The genus is based on holotype IVPP V11726, a crushed fossil found in 1998 at the Sihetun-locality. The layer it was discovered in, was argon-dated at an age of 124.6 million years. It was the first Chinese pterosaur fossil preserving the skull. It consists of the front half of a subadult, including a skull, lower jaws, pectoral girdle, sternum, wings, cervical and dorsal vertebrae, partial pelvis and metatarsals. The authors concluded that its slender hindfeet meant that it was forced to move quadrupedally on land, suggesting a piscivore lifestyle as a specialised soarer. [1]