When Shade and Marina were attempting to find Shade's colony in Silverwing, they encountered Zephyr in a cathedral spire. Despite his blindness, he is very knowledgeable and has incredible hearing, even being able to purportedly hear the stars. Zephyr tells them more of the mysterious bands, and their connection to Humans. He also provides Shade with limited information about his father, Cassiel, and shows Shade how to use his unique skill with sound. Eventually, the time comes for Shade and Marina to leave, and he bids them farewell.