Today, Spec pterosaur biota is dominated primarily by hastazhdarchine azhdarchids and nyctosaurids, with a few odd clades like carnocursorines and anatochasmines occuring here and there. All these forms belong to two main clades, Pteranodontia and Azhdarchidae, the pterosaurs that seemingly dominated terminal Cretaceous reccords on both worlds, Other pterosaurs, even the closely related clades, were thought to have been disappeared long in the poorly understood extinction events of the mid-Cretaceous, and as such to never have witnessed the start of the Mesozoic. Imagine the specologists' surprise, then, when they came across the exquisite lagerstattën of the Eocene, such as the Messel Pit. Here, while the familiar Maastrichtian pterosaurs were present, the dominant pterosaurs belonged to