Squigly is the last “surviving” member of the Contiello family, a long lineage of opera singers and among the Medicis’ most valued clients. Fourteen years ago, Squigly’s mother Selene obtained the Skull Heart, resulting in Lorenzo Medici ordering an attack on the Contiello family. Fraught with despair, Selene became the Skullgirl and revived her family as an undead army. What spared Squigly from becoming a mindless minion was the intervention of the Parasite Leviathan, the Contiello family’s friend and guardian.