"Tupandactylus imperator ."@en . . . . "(Unwin & Martill, 2007)"@en . . . "( )"@en . . "Tupandactylus"@en . . "* Ingridia"@en . "Synonyms"@en . . "Tupandactylus (meaning \"Tupan finger\", in reference to the Tupi thunder god) is a genus of tapejarid pterodactyloid pterosaur from the Early Cretaceous Santana Formation of Brazil. It is notable for its large cranial crest, composed partly of bone and partly of soft tissue. Tupandactylus is based on MCT 1622-R, an almost complete skull, found in the Crato Member of the Santana Formation. It was initially described as a species of Tapejara, but later research has indicated it warrants its own genus. The skull was toothless and had a prominent sagittal crest, only the base of which was bony: the front of the crest featured a tall bony rod extending up and back, and the rear of the crest had a long prong of bone projecting behind it. The bulk of the crest was made up of soft tissue similar to keratin, supported by the two bony struts."@en . "200"^^ . . . "Tupandactylus"@en . "Tupandactylus"@en . . . "Tupandactylus (meaning \"Tupan finger\", in reference to the Tupi thunder god) is a genus of tapejarid pterodactyloid pterosaur from the Early Cretaceous Santana Formation of Brazil. It is notable for its large cranial crest, composed partly of bone and partly of soft tissue."@en . . "* T. imperator"@en .